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Smooth Softshell Turtle
Apalone muticus muticus
by Jeff
LeClere
Status
The smooth softshell is listed as a Species
of Special Concern in Minnesota. This status is due to a lack
of records in the past.
Description
The smooth softshell is a large Minnesota
turtle and grows to a carapace length of 14 inches in females
(Conant and Collins, 1991 ). Males reach only half that length.
Softshells are most easily recognized by the appearance of
their carapace. It does not have the the bony scutes of our
other turtles. Instead, it is flat and leathery with very
flexible edges. It very much resembles a pancake. These turtles
differ from the very similar spiny softshells by having a
completely smooth carapace. The nose tapers to a point and
resembles a snorkle.There is NO ridge in each nostril, this
is also a good way to differ a specimen in hand from a spiny
softshell. Adult males are usually gray or brown with dots
and dashes on the back. Adult females are much more drab in
coloration and have large dark blotches on the carapace. Both
sexes have two light lines on the side of their head; one
behind the eye and one that starts under their snout. The
neck and feet are not patterned, but the sides of the head
may have a peach or pink wash. The feet of softshells have
extensive webbing--much more than any other Minnesota turtle
species. Their plastron is quite small and is white with no
distinctive markings. The young are about two inches at hatching
and are patterned like adult males.
Subspecies
The subspecies of smooth softshell found
in Minnesota is the midland smooth softshell, Apalone muticus.
Range
The smooth softshell is found in the Mississippi,
St. Croix, and Minnesota Rivers in Minnesota. Until recently
, there were very few records for this turtle and it was presumed
to be rare. The numerous new records for this species show
that it has an extensive Minnesota River Valley range. Although
nowhere near as abundant as the spiny softshell, Apalone
spinifera, smooth softshells can be expected to occur
in all of the major rivers in southern Minnesota.
Habitat
Smooth softshells are found in rivers,
streams, and, rarely, large lakes with sandy or muddy bottoms.
Sandbars are important for basking and egg laying sites.They
seem to prefer larger rivers and live in colonies along certain
portions.
Habits
These turtles are active from late April
and early May to late September or October. They bury themselves
in mud or sand at the bottoms of rivers or lakes to overwinter.
They are diurnal and sleep at night underwater buried in sand
or mud in the shallows or among submerged trees. They are
one of Minnesotas most aquatic turtle species. They bask on
sand or mud bars or on large logs and rocks, but always at
the waters edge. They are wary (more so than even spiny softshells)
and will quickly dash into the water at the slightest disturbance.They
are the fastest swimmers of Minnesotas turtles. Unlike spiny
softshells,these turtles are not aggressive and will remain
passive even when picked up.
Sometimes they rest buried in the sand
in shallow water. They extend their necks and use their long
noses as snorkels. They hunt during the day. Softshells can
remain submerged for up to five hours using either dermal
or cloacal respiration (Dunson, 1960). However, softshells
usually do not remain submerged for more than 20 minutes while
active (Stone et al, 1992b). They are capable of exchanging
gas through their skin in both water and air. The smooth skin
of soft shell turtles are three or four times as permeable
as other hard shelled turtles (Bentley and Schimdt-Nielson,
1970).
Smooth softshells breed in May and they
lay their eggs in June or July. Females usually dig nests
on sand bars that receive sunlight. This location is closer
to the water than Minnesotas other turtle species, so many
females are not observed (and killed) on their way to lay
the 12 to 30 eggs. Carnivorous mammals, however, dig up most
of the nests and eat the eggs. The brittle shelled eggs are
the shape of, but smaller than, ping-pong balls. The young
hatch in August or September.
Hatchlings overwintering in the nests
have not been observed in Minnesota. The young are patterned
like adult males, and have upturned noses and the sides of
the carapace droop covering the hind limbs and tail like a
blanket. Neonate softshells are eaten by a variety of carnivorous
vertebrates including large fish.
Food
Smooth softshells seem to prey mostly
upon insects, but other food such as worms, snails, clams,
isopods, crayfish, spiders, frogs, tadpoles, mudpuppies,fish,
young birds, small mammals, algae, and seeds have been recorded
(Ernst,Lovich, and Barbour, 1994). Males appear to consume
more terrestrial foods including cottonwood and mulberry seeds.
Females feed upon aquatic prey, mostly insect larvae (Plummer
and Farrar, 1981).
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